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A lot of it is because of their internal link structure. They link a lot of their pages together with "related videos" and whatnot which helps pass linkjuice from the front page. People also tend to link to tubesites a lot, but I really think it's more a factor of the number of pages and internal links.
Pornhub, for example, only has 2 million inbound links according to Yahoo. That's really not that many considering the amount of traffic the site gets. What they do have is 1.6 million internal pages that are all linked.
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